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sunfish (0.4). For Hotspot Pond, largemouth bass (2.0), western mosquitofish <br />(2.1), black crappie (1.2), green sunfish (0.4), and black bullhead (0.1). <br />The following conclusions from this research were: 1) the species <br />composition of both ponds was predominantly nonnative during all three sampling <br />periods, 2) native fishes used the two connected ponds mostly during runoff, 3) <br />five species of sub-adult and adult nonnative fishes moved between the two ponds <br />that were connected to the river; white sucker and largemouth bass migrated the <br />most between the two ponds. Roundtail chub was the only native fish to move <br />between the two ponds, 4) within four months following draining of Gardner Pond, <br />removal of nonnative fishes, and construction of a connection channel, nonnative <br />fishes invaded and recolonized the pond, and within eight months, five species <br />of nonnative fishes had successfully reproduced in Gardner Pond. Ponds in the <br />10-, 50-, and 100-year floodplain that have bottoms shaped and contoured as <br />depressions are havens that provide perennial, off-channel habitats for nonnative <br />fishes to grow and reproduce. Ponds that are depressions and that are connected <br />to the river allow sub-adult and adult fish uninhibited access to the river and <br />to other off-channel habitats connected to the river. Ponds with depressions <br />provide a constant supply source of nonnative fish that can potentially escape <br />to the mainstem river. While depressions benefit native fishes for a period only <br />during runoff, nonnative fishes appear to benefit more from such perennial, <br />lentic habitats, and 5) Hotspot Pond did not drain following runoff in 1996. If <br />ponds are reclaimed, steeper bottom slopes than that found in Hotspot Pond will <br />be required for ponds to drain during high-flow years to reduce or eliminate <br />perennial habitat for nonnative fishes. <br />Recommendations were to: 1) avoid connecting isolated ponds to the river <br />that are irregular- shaped, deep depressions because such habitats provide <br />perennial havens for nonnative fishes. If deep-depression habitats cannot be <br />rehabilitated by recontouring and reshaping to provide seasonal, ephemeral <br />habitat, or sloped to concentrate and harvest captive-reared fish, they should <br />be sufficiently diked to prevent escapement of nonnative fishes such as <br />centrarchids and ictalurids, and 2) back-fill gravel-pit ponds so that they are <br />sloped to flood and drain seasonally and reconnect them to the mainstem river to <br />benefit native fishes and to reduce proliferation of nonnative fishes. Such <br />modifications would allow the floodplain to function as an ephemeral wetland and <br />may be more cost-effective and practical for managing and controlling nonnative <br />fishes than long-term control measures such as screening, chemical treatment, or <br />mechanical pumping and removal of nonnative fish species. Whenever floodplain <br />habitats are connected to the river, hydrological access for fish should be by <br />surface connection. Underground culverts and pipes should be avoided because <br />some native fish might be shy and will not use such structures. <br />Alternative management options that should be considered by the Recovery <br />Program as habitat improvements to assist recovery of Colorado River endangered <br />fishes included: 1) coordinating Recovery Program efforts with the U. S. Army <br />Corps of Engineers, Colorado Mined Land Reclamation, and gravel-pit companies to <br />restore gravel-mining areas to function as historical terraces to benefit native <br />fishes, 2) notching dikes and levees protecting gravel pits by reconnecting <br />former floodplain habitats without deep depression ponds with the river to <br />enhance or restore the natural function of the river ecosystem. Dikes and levees <br />constructed to protect gravel-mining operations could be removed following gravel <br />x