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predation pressure was realized. It should be noted that had centrarchid numbers been <br />significantly reduced and YOY Colorado pikeminnow catch rates had increased, we <br />might hypothesize a cause-and-effect relationship, but drawing any such conclusions <br />would be difficult given the possibility for alternative explanations. <br />One potentially important observation was that green sunfish were often found <br />concentrated along shorelines containing rip-rap. Several backwaters were adjacent to <br />banks rip-rapped with large rocks or chunks of concrete and green sunfish seemed to <br />prefer the crevices associated with this material, particularly when other cover was <br />lacking. Submerged sticks in beaver dams or lodges served a similar function and <br />evidently provided enhanced habitat conditions for both green sunfish and largemouth <br />bass. Although little can be done to reduce beaver in the floodplain, construction of <br />dikes that contain rip-rap extending into the water should be discouraged. <br />In addition to structure, another important factor that may have enhanced <br />conditions for centrarchids during the project period was the lack of large flushing flows <br />through the side-channels. Spring flows were below average during 1998-2001 and <br />backwaters that might normally have stayed connected to the river became seasonally <br />isolated allowing for pond-like conditions to develop. Although most of these channels <br />flowed during spring, flows were apparently not high enough to flush fish, accumulated <br />sticks or rooted vegetation. <br />In addition to the high number of green sunfish and largemouth bass that were <br />found inhabiting upper Colorado River backwaters, there was also a surprisingly high <br />number of non-native black bullhead and white sucker. Bluegill, though not abundant, <br />were also found in higher numbers than anticipated. These high numbers of non-native <br />fish species, once thought rare, may have resulted from the use of a gear type not used in <br />past backwater sampling or may have resulted from real increases in these populations in <br />recent years, or perhaps resulted from a combination of these two factors. Channel <br />catfish, a ubiquitous member of the main-channel fish community, was found in <br />surprisingly low numbers in backwaters during late summer, fall, and early spring. <br />Based on degree of piscivory known from other studies (Osmundson 1987, <br />Nesler 2002) and relative abundance documented here, the greatest threat to survival of <br />22