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are formed. Deposits of cobble and gravel can also create islands resulting in multi-thread <br />channels or complex river reaches. Often associated with multi-channel sites are riffles at <br />the upstream end and sometimes pools at either the downstream end or to one side of the <br />riffle. Backwaters are formed at the downstream end of some side channels when inflow at <br />the upstream end ceases or is reduced as runoff flows subside. Eddies form at the interface <br />of the backwater mouth and the main channel. Thus, lateral movement of the channel <br />during high flow events is the process responsible for channel complexity and the creation <br />of preferred mesohabitats (Pitlick et al. 1999). <br />Adult Colorado pikeminnow in the Grand Valley prefer segments of river that <br />contain a diversity of habitat types. Partitioning the river into 0.65-km segments from <br />Loma to Palisade, Osmundson and Kaeding (1991) categorized segments as either simple <br />(single-thread) or complex (multi-thread or containing backwaters), and found that the river <br />consisted of about equal proportions of the two segment types. During radiotelemetry <br />studies, adults were located in complex segments 85% of the time during spring; 71%, <br />during summer; 62%, during winter. These complex segments are preferred presumably <br />because they contain more of the preferred mesohabitats (pools, eddies and backwaters) <br />than do simple segments (primarily consisting of fast and slow runs) and because a variety <br />of habitat types juxtaposed to one another allows more efficient exploitation of resources, <br />i.e., feeding and resting habitats are close together (Osmundson and Kaeding 1991). <br />Osmundson et al. (1995) selected four complex sub-reaches within the 15-mile reach <br />for habitat mapping during 1990-1991. The reaches selected were those heavily used by <br />adult Colorado pikeminnow. At a moderate base flow (1,630 cfs), backwaters comprised <br />5% of the total water surface area of the four sub-reaches; pools comprised 6%; eddies, <br />0.4%; riffles, 25%; slow runs, 56%; fast runs, 7%; rapids, 0.5%. Thus, even in complex <br />reaches, a small percentage of the total wetted channel area is comprised of non-run <br />habitats; the preference by adult Colorado pikeminnow for such sites underscores the <br />importance of creating and maintaining such features. <br />When flow is reduced and sediment input remains the same, fine-sediment <br />-deposition occurs in low velocity sites such as side channels, backwaters and the river <br />margin. Osmundson et al. (1995) and Van Steeter (1996) documented the deposition of <br />22