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containing optimum temperatures earlier in the season. Spawning migrations <br />and gonadal maturation of razorback suckers may largely be initiated by <br />photoperiod, with warm temperatures hastening final gonad maturation. In <br />the Grand Valley, sexual ripeness of razorbacks coincides with the period <br />of peak runoff (Fig. 13). Only during high flow conditions can razorbacks <br />of the upper basin find their optimum spawning temperatures in spring, but <br />they must leave the main channel to do so. The main channel during peak <br />flow averages 13.0 C (USGS gage data at Cameo; Appendix Table V). Razor- <br />back suckers can find water warmed to 20 C in off-channel habitats when <br />high spring flows flood low-lying areas adjacent to the river. These <br />still-water sites are warmed by direct sunlight and ambient air tempera- <br />tures and are generally much warmer than the main channel. Water tempera- <br />MAY 05-09 <br />MAY 10-14 <br />MAY 15-19 <br />MAY 20-24 <br />MAY 25-29 <br />MAY 30-03 <br />JUN 04-08 <br />JUN 09-13 <br />JUN 14-18 <br />JUN 19-23 <br />JUN 24-23 <br />JUN 29-03 <br />JUL 04-08 <br />0 5 10 15 20 25 30 35 <br />FREQUENCY M <br />Figure 13. Frequency of Colorado River peak flow dates during 1907-1989 <br />(N = 83 years) and capture dates during 1975-1988 of razorback suckers (N <br />42 fish) in spawning condition (expressible sex products present) in the <br />Grand Valley, Colorado. Two ripe razorback suckers captured in early April <br />1975 are not represented. Data from McAda and Wydoski (1980), Valdez et <br />al. (1982), USFWS (unpublished), and Osmundson and Kaeding (1989). <br />40