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ships. Other important variables, such as magnitude of summer flows and <br />temperature regimes are correlated with peak flows, and these in turn <br />could have a direct effect on larval production of some species or possi- <br />bly retainment of larvae within the study area (high summer flows might <br />facilitate long-distance transport of young). Thus, peak flows may be more <br />of an index to the type of water year rather than the controlling factor <br />itself. One possible direct-effect scenario for Colorado squawfish is that <br />some magnitude and duration of high flow may be necessary to prepare the <br />spawning substrate (Haynes et al. 1984). Though spawning may occur every <br />year, hatching success could vary depending on the degree to which the <br />cobble substrate is flushed of sediments. Results of studies on other <br />systems have demonstrated inverse relationships between accumulation of <br />fine sediment in fish spawning and rearing habitats versus fish survival <br />and abundance (Reiser et al. 1989). Studies are needed to determine the <br />importance of this variable for Colorado squawfish in the upper Colorado <br />River. Another possibility is that as numbers of non-natives increase in <br />years of low flow, predation-related mortality on squawfish might increase <br />(Valdez 7.990). <br />One might expect that numbers of larval Colorado squawfish transported out <br />of the reach via river currents would be higher in years of relatively <br />high summer flows, thus resulting in fewer YOY squawfish found in the <br />Grand Valley during fall of such years. Though flow effect on percent <br />larvae transported out of the reach is unknown, it is of interest to note <br />that total numbers of larvae and YOY were highest in 1986 despite,rela- <br />tively high summer flows that year (mean flow during August 1986 in the <br />18-mile reach downstream of the mouth of the Redlands power return, canal <br />23