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and periphyton (algae that grows on rock surfaces) in <br />support the native bluehead sucker (Catostomus disco <br />(Catostomus latipinnis). Upper reaches generally hav <br />downstream reaches; this serves to keep gravel and cc <br />Many aquatic insects require silt-free crevices around <br />support the native roundtail chub (Gila robusta), anot <br />Upstream reaches not only have higher food product: <br />the most suitable sites for spawning. As eggs deposi <br />drift with the current downstream. In the Colorado l <br />downstream of Moab, Utah having traveled a distanc <br />out and deposits of silt and sand provide many warm <br />of young, developing Colorado squawfish. During tl <br />backwaters and feed upon the zooplankton and chiro <br />become piscivorous, an abundance of small fishes coi <br />iese upper reaches; these algal types <br />As) and flannelmouth sucker <br />higher gradients than do more <br />ble substrates more free of silt and sand. <br />>cks for spaces to five; these insects in turn <br />,r forage fish of the Colorado squawfish. <br />n, but the clean cobble bars there provide <br />d in these areas hatch, the resulting larvae <br />ver, most of these larvae end up <br />of 60-180 miles. There the river flattens <br />sites ideally suited to the rearing <br />first year of fife, squawfish seek out these <br />iid larvae found there. Later, as they <br />ue to provide them with ample forage. <br />As Colorado squawfish become larger and approach ulthood, larger forage fish are required to <br />maintain growth rates. However, larger-sized forage sh are scarce in these downstream reaches <br />and maturing Colorado squawfish begin to move up d down the river seeking better feeding <br />grounds (Fig. 3). Eventually, the abundant supplies o native fish in the upper river are discovered <br />and adult squawfish remain in these upper reaches the remainder of their lives. As a result, the <br />average size of Colorado squawfish increases as one moves up river, with the greatest numbers of <br />large fish found above Westwater Canyon (Fig. 4). Even above Westwater, the differences in <br />average size among the three river strata are striking. Most Colorado squawfish in the Grand <br />Valley are clearly larger than those in the Loma-to-W stwater reach, and within the Grand Valley, <br />the 15-mile reach contains the greatest proportion of 1 ge, adult fish. <br />The complete life cycle.of the razorback sucker, in 1 <br />largely unknown but may be similar to that of Cola <br />concentrated in upstream regions of the Green and <br />sites in the upper Green River have also been found <br />evidence suggests that bottomlands up and down be <br />for larvae when these habitats flooded during the fo <br />as of its river-wide habitat use pattern, is <br />squawfish because adults are generally <br />orado rivers and larvae from spawning <br />drift long distances down- river. Other <br />rivers historically served as nursery areas <br />;rly higher spring runoff period. <br />To understand the habitat needs of these large-river <br />takes into account all phases in their respective life c <br />particular species, population dynamics occur at the <br />importance of a particular area, like the 15-mile reac <br />must be viewed in the context of how it fits into the <br />habitat cannot support a viable population just as a i <br />cannot. Without taking the above discussion into ac <br />erroneous view of the importance of the 15-mile rea <br />the 185 miles of the Colorado River (above the Gre, <br />important in helping maintain populations of Colora <br />I, an ecological perspective is required that <br />-les and a recognition that for these <br />ndscape scale. Thus, the relative <br />cannot be viewed in isolation; instead, it <br />rger picture. A river without good adult <br />er with poor spawning or nursery habitat <br />)unt, one might arrive at a simplistic and <br />G that is, because it constitutes only 81/6 of <br />River confluence), it is probably not terribly <br />squawfish and razorback sucker. <br />4