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1.3 RIPARIAN VEGETATION OF THE FLOOD- <br />PLAIN <br />Throughout the desert Southwest, a <br />few plants are uniquely adapted to the <br />floodplains of seasonally fluctuating <br />streams. These riparian plants exist <br />where their roots are in the capillary <br />fringe of the water table. They may <br />extend only as far from the channel as <br />the stream exerts its influence <br />through a water table. This strip of <br />vegetation Is often used to define the <br />floodplain of a river, and creates a <br />marked contrast as a ribbon of green <br />bisecting the desert uplands (Figure <br />6). <br />WETLAND <br />z Mes uite Cottonwood-willow Mesquite replacing <br />z woo land forest cotton woo d?w i 11 o w z <br />a mar sr?snA <br />o <br />F m c <br />m U m <br />?j Q « U <br />Flood old river Flow channel Flood terrace <br />terrace channel <br />High water channel <br />FLOODPLAIN <br />Figure 6. Semidiagrammatic represen- <br />tation of riparian communities in warm <br />temperate to subtropical habitats of <br />the American Southwest. Adapted from <br />Minckley and Brown (1982). <br />The natural vegetation associa- <br />tions along the lower Colorado River <br />were well described by Grinnell <br />(1914). Belts of riparian vegetation <br />stretched for many kilometers and <br />filled the broad alluvial valleys. <br />The dominant riparian forest species <br />were cottonwood (Po°?ulu_s fremontii) <br />and black or Goodding willow (Salix <br />gooddinavil). These occurred primarily <br />on the "first bottom" and braided <br />channels, in association with the <br />understory shrub seepwillow (Baccharis <br />sal icifolia [B. glutinosa]) and <br />occasionally other willows (e.g., <br />coyote willow, Salix extgua) and emer- <br />gent species (e.g., cattail, [T=ha <br />spp.] and bulrush [Scirgus spp.]) <br />(Figure 7). As an adaptation to a <br />frequently flooded environment, these <br />plants were fast-growing and relative- <br />ly short-lived. In fact, their exis- <br />tence was ultimately dependent on the <br />cycle of annual floods that created <br />new silt beds for seed germination. <br />However, these and other native plants <br />cannot tolerate prolonged inundation. <br />Where such long-term flooding <br />persisted, such as in oxbow lakes, <br />emergent marsh vegetation became <br />established (Figure 8). This marsh <br />vegetation consisted of cattails, <br />bulrushes or tules, and in the <br />southern portions of the valley, cane <br />( ragmites australis [P. communis]). <br />Along the drier sites adjacent to <br />the willow and cottonwood stands a <br />shrub, arrowweed (Tessaria sericea <br />[Pluchea sericea]), often formed dense <br />monotypic belts or small strands in <br />some areas (Figure 9). Where the <br />floodplain of the first bottom escaped <br />inundation for a number of years, the <br />rare screwbean mesquite (Prosopis <br />pubescens) grew in association with <br />willows (Figure 10). <br />A very different type of riparian <br />vegetation occurred on the second <br />bottoms than that which existed ad- <br />jacent to the river. The dominant <br />tree in the second bottom was honey <br />mesquite (erosoRL gl andu loss [P.. <br />juliflora]) (Figure 11). This tree <br />formed relatively sparse monotypic <br />woodlands. The long roots of the <br />mesquite must find permanently moist <br />soils to ensure survival, yet the tree <br />itself apparently cannot tolerate <br />inundations of even a few weeks. In <br />addition to honey mesquite, several <br />9