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<br />Identifying and interpreting 1) the timing of organic <br />inputs and patterning of natural quality gradients to <br />reaches along the river ecosystem, 2) strategic patterns <br />adopted by consumer assemblages for processing organic <br />matter available to the river system, and 3) process <br />efficiencies associated with the utilization of energy <br />throughout the continuum (p. 1). <br />9 <br />According to the theory, river systems exhibit heterotrophy i <br />4 <br />headwaters where shading and organic input are high, autotrop,y in <br />mid reaches where insolation and temperatures increase, and h tero- <br />trophy in lower reaches where increases in suspended material cause <br />reduced light penetration. <br />No works have specifically addressed the River Continuu Theory <br />since its proposal although a wide range of studies have previously <br />been done which address key elements of the theory and lend s pport <br />to its relevance. <br />. Fisher and Likens (1973) found that up to 99 percent of t e <br />organic energy in a low order stream was of allochthonous orig'n. <br />Minshall (1967) found similar heterotrophic conditions in a to order <br />stream but suggested that primary production played a more imp rtant <br />role in larger streams. Cummins discusses various aspects of he <br />theory in a 1977 work. <br />. The transitions between heterotrophy and autotrophy are <br />controlled by a host of interacting factors including discharg rates <br />and velocities, temperature, insolation, and organic and inorg nic <br />. inputs (Cummins 1974). Thomas and O'Connell (1966) showed tha <br />temperature and insolation are primary factors affecting Bros <br />production but suggested that a combination of other factors, <br />primarily allochthonous materials, control the net amount of p imary <br />0