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<br />I <br />in the Maybell reach, 49 northern pike in the Lily Park reach, and 1 in Yampa Canyon. Sizes <br />of northern pike examined ranged from 87 to 1036 mm total length and examination of the <br />length frequency distribution of these fish shows this sample was representative of the sample <br />population size distribution (Figure 11). Also, the size distribution of northern pike containing <br />food in their gut was quite similar to the size distribution of northern pike with empty stomachs. <br />Overall, 63.8 % of the northern pike examined had ingested fish prey. Non-fish food <br />items were found in 7.5 % of the northern pike examined and 36.1 % of the northern pike had <br />no food in the stomach (Figure 12). Twelve species of fish prey ingested by northern pike were <br />identified. In order of abundance, fish prey included redside shiner Richardsonius balteatus, <br />roundtail chub Gila robusta, white sucker Catostomus commersoni, speckled dace Rhinichthys <br />osculus, flannehnouth sucker Catostomus ladpiwds, mottled sculpin Cottus bairdi, bluehead <br />sucker Catostomus discobolus, plains killifish Fundulus zebrinus, fathead minnow Pimephales <br />promelas, common carp Cyprinus carpio, green sunfish Lepomis cyanellus, and sand shiner <br />Notropis stramineus. Northern pike prey composition was dominated by four species; two non- <br />native species comprised 42.3 % of the fish eaten and two native species comprised 13.6 %. By <br />species, redside shiner comprised 33.3 %, roundtail chub comprised 9.5 %, white sucker <br />comprised 9.0 %, and speckled dace comprised 4.1 % of the prey composition. The remaining <br />seven species made up 9.5 % of the prey identified and 34.7 % of the fish prey could not be <br />identified. Only in two instances were two different fish species found together in a northern <br />pike stomach. In the spring months of April through June, 60% of the 160 northern pike <br />stomachs contained food. In the fall months of September through November, 74% of the 31 <br />northern pike stomachs contained food. Other food items identified in northern pike stomachs <br />included Cladocera, Annelida, Insecta, Decapoda, and Anura. <br />In two other studies occurring downstream in the middle Green River below the <br />confluence with the Yampa, Tyus and Beard (1990) found 54.5 % of 58 northern pike stomachs <br />to be empty. Frequency of occurrence of fish species ingested were red shiner Cyprinella <br />lutrensis at 7.3 %, followed by fathead minnow, flannehnouth sucker, and speckled dace. <br />Another eight fish species comprised 13.6 % of the fish prey and 10.6 % could not be identified. <br />Non-fish prey occurred in 2.4% of the northern pike. Overall, nine introduced fish species <br />comprised 25% of northern pike stomach contents and three native species comprised 9%. <br />Crowl et al. (1995) examined 26 northern pike and found stomach contents comprised of 5 % <br />juvenile Colorado squawfish and 72% other cyprinid species, followed by mice, crayfish and <br />sucker species. Seasonal differences were also noted by Crowl et al. with 89 % of the northern <br />pike containing food in the spring and 12.5% in the fall. In studies elsewhere, Rich (1992) <br />observed empty stomachs in 63-74 % of northern pike during the spring and 45-69 % in the fall. <br />Headrick (1985) also noted high percentages of empty stomachs in northern pike. Northern pike <br />feeding was determined to be more intense during spring and fall (Seaburg and Moyle 1964, <br />Weithman and Anderson 1977). Headrick and Carlin (1993) noted condition factors for <br />northern pike in impoundments was highest in the spring and lowest in late summer. They <br />found northern pike would lose weight during the summer and gain again in the fall. <br /> <br />DRAFT <br />March 12, 1996 <br />35 <br />1