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INTRODUCTION <br />This study represents an evaluation of the present state of knowl- <br />edge concerning the aquatic flora and fauna of the upper Colorado River <br />basin. Relative detail of treatment varies according to the quantity <br />and quality of available information, and all the information that would <br />be desirable is not available for any particular subject area. Emphasis <br />of the report is placed on the endangered and threatened fishes, their <br />status, habitat requirements, and overall ecology. <br />The upper Colorado basin comprises an area of approximately 109,580 <br />square miles, encompassing all of the Colorado River drainage above <br />Lee's Ferry, Arizona. Streams in this basin begin as clear, cold, <br />first-order streams draining the mountains of Wyoming, Utah, and Colo- <br />rado. They join to form larger streams that flow into the desert areas <br />of southeastern Utah, western Colorado, and northern Arizona and New <br />Mexico. In these regions they are large third- to fourth-order rivers <br />that flow through some of the world's most spectacular canyons. <br />The river system of the upper basin is geologically old and has <br />taken millions of years to carve its way through thousands of feet of <br />rock. Headwater streams typically flow over igneous and metamorphic <br />rocks. Sedimentary rocks predominate in and below areas of second- or <br />third-order streams and in some headwater regions. The aquatic fauna of <br />this system are also very old, for they evolved in accord with the <br />geological changes that took place in the basin. The indigenous native <br />fish species of the system have been-isolated from other drainages <br />nearly as long as the present Colorado drainage has existed, and have <br />largely evolved independently of fishes of other regions. This has <br />resulted in a high degree of endemism in Colorado River fish species. <br />Other aquatic flora and fauna have terrestrial dispersion mechanisms <br />which have prevented the development of a high degree of endemism. <br />The Colorado River basin has historically been a river of great <br />fluctuations in flow, turbidities, and temperature. However, man- <br />induced changes have dramatically influenced these regimes. Man's <br />activities within the basin have increased sediment load and total <br />dissolved solids (TDS). Man-made reservoirs in turn precipitate out <br />this load, resulting in drastically reduced turbidities and flow, along <br />1