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Acknowledgments <br />This study was funded in part by U. S. Fish and Wildlife Service, <br />Bureau of Reclamation, National Park Service, and the Northern Colorado <br />Water Conservancy District. J. Beard, P. Clevenger, and L. Trinca were <br />among several who assisted with data collection. W. C. Starnes and two <br />anonymous reviewers improved an earlier draft of the manuscript. <br />17