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in 1956, exports water from the Colorado River <br />headwaters to the South Platte drainage in eastern <br />Colorado (Pennak 1963). Water is stored in Willow <br />Creek Reservoir, Lake Granby, Shadow Mountain <br />Lake, and Grand Lake and is delivered to the <br />eastern slope through the 21-km Adams Tunnel for <br />flood control, irrigation, municipal supplies, <br />hydroelectric power, and recreational facilities. <br />The Roaring Fork River joins the Colorado River <br />at Glenwood Springs, Colorado. The largest <br />tributary of the Upper Main-stem Colorado, the <br />Gunnison River, enters the Colorado at Grand Junc- <br />tion, Colorado. In Utah, the Dolores River enters the <br />Colorado; it is the last major tributary upstream of <br />the Colorado-Green River confluence. <br />The San Juan Sub-basin <br />The San Juan Sub-basin is the drainage between <br />the junction of the Green and Colorado rivers and <br />the Lower Colorado River Basin. Its 99,200 km' area <br />is in Colorado, Utah, New Mexico, and Arizona. <br />The Colorado River below the mouth of the Green <br />River passes through Cataract Canyon and remains <br />entrenched in a deep canyon as it flows about 200 <br />km southwestward to Lee Ferry (Frontis.). Much of <br />the stretch of river below Cataract Canyon is now <br />part of Lake Powell. Glen Canyon Dam and Reser- <br />voir (Lake Powell) were authorized by the Colorado <br />River Storage Project Act in 1956; the dam, a few <br />miles below the Utah-Arizona line, was completed in <br />1964 (Upper Colorado River Commission 1980). <br />Principal Colorado River tributaries which now <br />enter Lake Powell are the Dirty Devil, Escalante, <br />and San Juan rivers. <br />The San Juan River arises on the southern slopes <br />of the San Juan Mountains in southwestern Col- <br />orado, flows southwestward into New Mexico, and <br />then turns back into the southwestern corner of <br />Colorado before entering Utah. It flows through a <br />deep canyon before entering Lake Powell. The <br />Navajo Storage Unit of the Colorado River Storage <br />Project was completed in 1963 to regulate San Juan <br />River flows for irrigation and municipal purposes <br />(Upper Colorado River Commission 1980). <br />The Paria River joins the Colorado River about 25 <br />km below Glen Canyon Dam and 1.6 km north of Lee <br />Ferry. <br />Dams, Reservoirs, and Water Diversions <br />We have not attempted to mention all dams, <br />reservoirs, and water diversions in the Upper <br />Colorado River Basin. Spofford (1980) referred to <br />nine principal federal reservoirs in the basin. A com- <br />plete list of the 21 participating projects of the <br />Colorado River Storage Project authorized by Con- <br />gress appeared in the Thirtieth Annual Report of <br />the Upper Colorado River Commission (1978). Some <br />of these are still in planning phases or under con- <br />struction. Other, non-federal, projects such as the <br />Moffat and Roberts tunnels of the Denver Water <br />Board also exist. A complete list of existing and <br />planned projects is very difficult to develop and <br />keep up-to-date. <br />Many diversions, like the Colorado-Big Thompson <br />Project, result in export of water from the Colorado <br />River Basin. Water diverted from the Upper Col- <br />orado River Basin is used in the Arkansas River, <br />Platte River, and Rio Grande River basins and in the <br />Great Basin. Martin (1981) reported, for example, <br />that 27% of Colorado's legal share of Colorado River <br />water is diverted to eastern-slope cities from Fort <br />Collins south to Pueblo and that water needs of <br />these cities are expected to increase by at least <br />200% in the next 30 years. Schad (1980) noted the <br />irony in the Colorado River Basin, which drains <br />some of the nation's more and lands and has the <br />lowest run-off per square mile of any major river <br />basin, being the source of such a large number of in- <br />terbasin transfers. <br />Conditions in the Basin <br />Iorns et al (1965), Joseph et aL (1977), and Bishop <br />and Porcella (1980) summarized conditions in the Up- <br />per Colorado River Basin. A broad range of climatic <br />and streamflow conditions exist in the basin; annual <br />precipitation varies from over 127 cm in mountains <br />to under 15 cm in desert areas. <br />Seasonal streamflow is derived primarily from <br />snowmelt in mountainous areas, and historic unit <br />discharge rates decrease rapidly as tributary <br />streams flow from their headwaters into less humid <br />areas (Bishop and Porcella 1980). Significant varia- <br />tions in annual discharge have occurred from year to <br />year and over periods of years (due to long-term <br />climatic trends). Progressive 10-year running <br />averages of estimated "virgin flow" (if the stream <br />were in its natural state and unaffected by the ac- <br />tivities of man) at Lee Ferry have ranged from 16.0 <br />to 17.8 billion m' since 1970, and the 1896-1980 long- <br />term annual average virgin flow at Lee Ferry is <br />about 18.3 billion m' (Upper Colorado River Commis- <br />sion 1980). The Upper Main-stem Colorado Sub-basin <br />contributes the greatest volume of water and the <br />San Juan Sub-basin the least. Joseph et aL (1977) <br />and Spofford (1980) summarized flow data from <br />selected U.S. Geological Survey gaging stations in <br />the Upper Colorado River Basin. <br />Joseph et aL (1977) recognized three distinct <br />stream zones in the Upper Colorado River System. <br />Their upper (headwater) zone was characterized by <br />cold, clear water, high gradient, and rocky or gravel- <br />ly substrate and was regarded as ideal habitat for <br />cold-water fishes. In this zone primary production <br />(mainly by "periphytic" algae) was considered <br />"significant" and benthic invertebrate production <br />"substantial". An intermediate zone occurs as <br />streams flow from the upper zone; there, water <br />warms, discharge increases, waters are turbid dur- <br />2