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northern lakes {Serns 1982, 1983). Few studies have related electrofishing CPE to population <br />density of fish in rivers, however Paragamian (1989) found that electrofishing CPE of <br />smallmouth bass M. dolomieui in rivers was significantly greater during nighttime than it was <br />during daytime. <br />In the upper Colorado River basin, mean CPE was poorly con elated with independent <br />population estimates of Colorado pikeminnow in most cases. However, estimates made by <br />Nesler (2000) for the Green and White rivers were correlated with ISMP catch rates for <br />1992-1997; no correlations were found for the Colorado and Yampa rivers. The correlations <br />occurred in the two rivers with the greatest increase in mean CPE during the study period. <br />Nesler's (2000) estimates were made using capture-recapture data collected during all studies <br />funded by the Recovery Program. The data were collected with a variety of sampling designs <br />and capture records from ISMP comprised a substantial portion of those data. However, the <br />data were not collected using amulti-pass sampling methodology specifically designed to <br />develop amark-recapture population estimate (e.g. Pollock et al. 1990). <br />Osmundson and Burnham (1998) and Osmundson (2002) estimated population size of <br />Colorado pikeminnow in the Colorado River during two periods encompassed by ISMP - <br />1991-1994 and 1998-2000 - using a multi-pass sampling design. ISMP data were included <br />as portions of one pass in their studies, but considerable data were collected specific to the <br />study. They noted a consistent increase in population size in the Colorado River which <br />roughly corresponded with the increase in CPE recorded by ISMP. However, the two values <br />were not correlated. Although CPE in the Colorado River increased from 1986-1998, it was <br />relatively stable (~1 CPM/hr) during the years when population estimates were available. In <br />addition, estimated population size continued to increase through 2000 (although not <br />significantly) while CPE decreased during the last two years of ISMP. Osmundson (2002) <br />also noted a decline in CPE (measured as CPM per trammel net set} in the later years of his <br />study, even though the annual estimates increased slightly. <br />Osmundson's primary collection technique involved trapping Colorado pikeminnow in <br />backwaters using trammel nets (scare and snare; described in Osmundson and Burnham <br />1998); shoreline electrofishing was used to supplement the scare-and-snare data in reaches <br />without sufficient backwaters. Scare and snare is more efficient than electrofishing because <br />Colorado pikeminnow are attracted to backwaters during runoff and are more susceptible to <br />capture. The different sampling technique may be partly responsible for the lack of <br />correlation between ISMP data and the Colorado River population estimates. CPE should be <br />related to population size in some manner, but the many variables associated with sampling in <br />difficult environments such as the rivers of the upper basin suggest that the nature of the <br />relationship will be difficult to identify. <br />19 <br />