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ACKNOWLEDGMENTS <br />Thad Bingham, Tom Chart, Brian Scheer, Andrew Scott, Rick Smaniotto, Matt Toner, <br />Mike Klopf, and Sara Hollerich assisted with field work. Ron Ryel used program CAPTURE <br />to produce population estimates. Reviews by Tom Czapla, Bruce Haines, John Hayes, and <br />Richard Valdez improved earlier versions of this report. <br />This study was funded by the Upper Colorado River Endangered Fish Recovery <br />Program. The Recovery Program is a joint effort of the Fish and Wildlife Service, Bureau of <br />Reclamation, National Park Service, Western Area Power Administration, states of Colorado, <br />Utah, and Wyoming, Colorado Energy Distributors Association, Upper Basin water users, and <br />environmental organizations. <br />iii <br />