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Winter 2 with low flow). The Yampa River was ice-covered from early December to early March <br />of both winters. <br />The ten Green River fish in Winter 1 (ice-&ee conditions) were observed in runs, slackwaters, <br />eddies, and backwaters for 43, 39, 12, and 6 percent, respectively, of the time observed with <br />radiotelemetry. In Winter 2, nine different Green River fish in ice-free conditions were observed in <br />runs, slackwaters, backwaters, and eddies for 64, 29, 5 and 2 percent, respectively, of the time. The <br />same nine fish in ice-covered conditions occupied runs, backwaters, and slackwaters for 69, 17, and <br />14 percent, respectively, of the time observed. Greater use of runs in ice-free conditions in Winter <br />2 was attributed to lower main channel velocities from lower river flows. Greater use of backwaters <br />under ice in Winter 2 was attributed to greater availability of these off-channel habitats and use of <br />ice as cover by foraging fish. <br />Valdez and Masslich (1989) hypothesized that use of these habitats reflected resting and <br />feeding activities. Overwintering adult Colorado squawfish appeared to rest in slow midchannel runs <br />and slackwaters, and fed in eddies and backwaters. Increased use of backwaters under ice-covered <br />conditions in Winter 2 indicated use of ice as cover during foraging. <br />The Green River fish used habitats with low average channel velocity of 0.0 to 0.3 m/sec and <br />average depth of 0.75 to 1.35 m (Table 4). Fish in midchannel runs were commonly in deep sand <br />troughs, while fish in slackwaters were immediately downstream of sand shoals, at the base of steep <br />sand banks. Fish in rocky areas remained in eddies behind rock jetties, but did not use talus or <br />nearby vegetated shorelines (primarily tamarisk). <br />The Yampa River fish also used distinctly different ovecwintering habitat in the two years <br />observed. During Winter 1, the fish most often used off-channel habitats (i.e., backwaters and <br />embayments), while the fish in Winter 2 used primarily main-channel habitats (runs, eddies, and <br />pools). The difference was attributed to flows higher than the historic mean in Winter 1 and lower <br />that the historic mean in Winter 2 (Figure 11). High use of pool habitat in Winter 2 was also <br />attributed to incorporation of spool-dominated reach (Lily Park) into the study area in the second <br />year of the study. <br />The Yampa River fish seemed to use habitat according to availability. When a variety of <br />habitats was available--during high .flows in Winter 1--fish selected off-channel habitats over main <br />channel types. In Winter 2--whea flow was low--many off-channel habitats were desiccated or ice- <br />filled and not available to the fish. Wick and Hawkins (1989) noted that the fish used eddies and <br />pools in areas devoid of backwaters and embayments, such as at Lily Park But, in areas with a <br />27 <br />