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INTRODUCTION <br />The Yampa River originates on the western slope of the Rocky <br />Mountains near Yampa, Colorado. It is the largest tributary to the • <br />Green River. The Yampa flows first in a northerly and then in a <br />westerly direction to its confluence with the Green River in Dinosaur <br />National Monument (Fig. 1). The Yampa River can be characterized as a <br />system of extreme hydrologic events. Dramatic seasonal fluctuations in <br />flow create equally dramatic changes in temperature, turbidity, <br />conductivity, and quantity of allochthonous material. These seasonal <br />variations in flow and water quality also exhibit long-term pulses <br />caused by annual variations in precipitation and accumulation of snow in <br />mountainous regions of the river watershed. Amid these hydrologic <br />extremes the endemic Colorado squawfish evolved a unique migratory <br />reproductive strategy. <br />Historically, migration of Colorado squawfish (Ptychocheilus <br />lucius) was considered a predictable summer event in many of the rivers <br />of western Colorado (Sigler and Miller, 1963; Joseph et al., 1977; <br />Behnke and Benson, 1980). However, neither origin nor destination of <br />the migrants was documented. It was assumed that the migration was <br />related to spawning. <br />Recent studies, conducted by the U.S. Fish and Wildlife Service <br />(USFWS), Colorado Division of Wildlife (CDOW), and Utah Division of <br />Wildlife Resources in the Upper Colorado River basin (Wick et al., 1981; <br />e <br />Haynes and Muth, 1982; Miller et al., 1982a,b, 1983b; Tyus et al., <br />1982a,b; Radant et al., 1983) produced new data regarding the movements <br />1