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connectivity to the river floodplain. Energy dynamics and food production are <br />strongly dependent on floodplain habitats in which shallow littoral zones <br />containing rich organic matter yield much higher densities of zooplankton than <br />habitats in river channels (Hynes 1970; Welcomme 1985, 1989). <br />l Zooplankton densities in Ouray National Wildlife Refuge floodplain habitats <br />were 300 organisms/liter, 10 times those found in adjacent river backwaters <br />during spring (Mabey and Shiozawa 1993). <br />Razorback sucker spawning observations <br />On the middle Green River, razorback sucker congregate at the <br />spawning bars just prior to peak flows about 16 kilometers (km) upstream from <br />Jensen, Utah (Figure 4) and on the lower Yampa River 0.8 kin above its <br />confluence with the Green River (Tyus and Karp 1990). The Green River <br />spawning site located near river kilometer 500 appears to be the most heavily <br />used with over 71% of all captures of razorback sucker in spawning condition <br />from the Green River being from this location (USFWS data base, Vernal <br />Field Station, B. Haines, pers. comm. 1997) Other spawning sites exist, such <br />as the small cobble bar located at river km 492, but size may limit larval <br />1 production (Figure 5a). <br />"J In the middle Green River spawning usually begins after spring flows <br />start but before spring peak flows. Temperature averages 14 C (range 9-17 C) <br />through the spawning period (Tyus and Karp 1990). Ripe adults were <br />collected over coarse sand substrates and in the vicinity of gravel or cobble <br />bars with an average water depth of 0.63 m. (Tyus and Karp 1990). Average <br />velocity over the bars was 0.74 in s''. On the Yampa River spawning site <br />6. a <br />velocity associated with spawning fish is 0.3 m s'' at depths of 0.7-1.0 in <br />(McAda and Wydowski 1980). Direct observation of spawning behavior on the <br />middle Green River sites was not made because of high turbidity (Tyus 1987). <br />Razorback sucker females with anal fins almost completely worn off have been <br />12