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<br />an overall increase in the density of invertebrates. The number of stoneflies <br />and caddisflies has decreased dramatically at this site since impoundment, <br />while the densities of mayflies and dipterans appear to have increased <br />substantially. Species collected in pre-impoundment studies, but not <br />in this study, are listed in Table 7. <br /> <br />Biomass measurements are not directly comparable since we used volumetric <br />analyses in order to preserve the specimens. The percent composition <br />can be used for comparative purposes. The trend is the same as the density <br />trend; a very marked decrease in stonef1ies and caddisflies 2nd a six- <br />fold increase in mayflies and 2.5-fo1d increase in dipterdns. <br /> <br />Comparisons of 1970 and 1980 Data on the Fisher River <br /> <br />Comparison of our data with a 1970 study of the Fisher River (May <br />1972) indicates that there have been similar changes in the relative <br />percent composition by density and biomass of insect orders in the Fisher <br />River in the last 10 years. A comparison of the percent composition <br />of annual means in density and biomass at two stations from the 1970 <br />study and from our study site on the Fisher River (Table 8) indicates <br />a decrease in the importance of stoneflies and caddisflies and an increase <br />in dipterans. This is likely due to long-term changes associated with <br />channelization of the Fisher River at the time Libby Dam was constructed, <br />and other changes associated with logging or agriculture in the Fisher <br />River drainage. <br /> <br />Comparisons of Present Populations of Invertebrates in Kootenai and <br />Fisher Rivers <br /> <br />A notable feature of the benthos in the Kootenai River was the paucity <br />of stonef1ies which generally comprise less than 0.1 percent of the density <br />of benthic invertebrates (Table 4). There were no species of stonefly <br />common in the Kootenai River as opposed to about 14 species which were <br />common in the Fisher River. Seventeen species of stonef1ies were collected <br />in the Kootenai River compared to 23 species in the Fisher River and <br />42 species in the partially regulated Flathead River. Their absence <br />may be related to changes in the substrate, to higher fall and winter <br />water temperatures, or to availability of oxygen (see later section of <br />possible problems with low dissolved oxygen). <br /> <br />Shifts in the species composition of mayflies were found with increasin9 <br />distance downstream from the dam and between regulated and control sites <br />(Tables 9 and 10). Two species of mayflies (Ephem~etla in~~ and <br />Ba~ tnieaudat~) predominate near the dam. They are species with <br />several generations per year, and apparently are prolific enough to with- <br />stand population losses due to frequent stranding and downstream drift <br />caused by flow fluctuations. The heptageniid mayflies (Epeo~~, Rhit~ogena) <br />were found in very low numbers near the dam, but increased at the downstream <br />stations. They were far more abundant in the Fisher River than in the <br />Kootenai River (Table 10). Rapid water fluctuations and increased algal <br />growths probably impair the efficiency with which they can maintain their <br /> <br />_?1_ <br />