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<br />Razorback sucker <br /> <br />Spawning of razorback sucker, like Colorado squawfish and humpback chub <br />is associated with changes in the hydrograph. Razorback sucker spawn on <br />the ascending limb of the hydrograph (Figure 7, from Tyus and Karp 1989). <br />As with the Colorado squawfish, razorback sucker are known to use n'llo <br />primary spawning areas, one on the Green River near RM 311 and the other <br />in the Yampa River just above the confluence with the Green River. <br />Individuals have been observed to move as far as 100 river miles to a <br />specific spawning site (Tyus and Karp 1990, and Modde et al. in <br />preparation). Movement to the spawning bar appeared more related to <br />discharge rate than temperature (Figure 8). As with Colorado squawfish, the <br />pattern of the hydrograph rather than absolute flows tend to initiate <br />movement to spawning locations. However, in 1995 fish implanted with radio <br />transmitters were observed to move to the spawning locations in late April, <br />but, larvae were not collected downstream until June when water" <br />temperatures exceeded 140 C (Fish and Wildlife Service unpublished data). <br />Thus, although adults seem to be attracted to the spawning sites by <br />discharge, spawning may not occur until temperatures reach approximately <br />140 C (Tyus and Karp 1990). <br /> <br />In general, the spawning requirements of all three listed species, known to <br />reproduce it the Yampa River is determined by the combination of flows and <br />temperature, both of which are related. The spawning activities of these fish were <br />influenced more by the pattern of flow: Le. rise or drop in the hydrograph, in any given <br />year than a specific magnitude of flow. Each species migrates and spawns at a <br />different stage of the hydrograph (Figure 9). Therefore, the occurrence of a natural <br />hydrograph is an important feature to the reproductive success of endangered fishes <br />and represents a rationale as to why the largest natural populations of Colorado <br />squawfish (Tyus 1991) and razorback sucker (Lanigan and Tyus 1989, Modde et al. <br />accepted for publication) occur in waters influenced by the Yampa River hydrograph. <br />It appears that temperature and the shape of the hydrograph have a greater influence <br />on behavior relating to the initiation of spawning activity than the magnitude of the <br />hydrograph. However, it is the magnitude of the flows which influence the <br />geomorphology of the river and effect habitat availability. Thus, although fish appear <br />to cue to specific patterns of the hydrograph, the magnitude of the hydrograph is <br />responsible for creating available spawning, nursery and adult habitat. <br /> <br />Association of Nonnative Fishes with Native Fishes <br /> <br />Native fishes were reported to be more abundant than nonnative fishes in the <br />canyon reaches of the Yampa River than in either the upper (RM > 54 or lower <br />reaches (RM < 1.8) (Figure 10. Appendix 2). Among all Yampa Canyon (RM 1.8 to <br /> <br />14 <br />