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<br />~) <br />'.1 <br />. -.J <br /> <br />Chapter 1 <br />INTRODUCTION <br /> <br />"01 <br />.. ~.1 <br /> <br />'.1'1. <br />rB~ <br />j <br />~ <br />] <br />..I <br /> <br />::j.. <br />;;,: <br /> <br />] <br />,:.1 <br /> <br />." <br /> <br /> <br /> <br />....J <br />.. <br /> <br />~.' <br /> <br /> <br /> <br />~:: <br />,. <br />.. <br /> <br />j <br />1 <br /> <br />Native fishes of the upper Colorado River now experience a very <br />different environment than the conditions under which they evolved. As <br />early as 1000 A.D., water from the Colorado River was removed for <br />irrigated agriculture (Graf, 1985). Diversions were small and had no <br />significant effects on the ecology or channel morphology of the river, but <br />in the late 1800s this began to change. The first major water development <br />on the upper Colorado River was the Grand Valley diversion, completed <br />in 1883. With a capacity of 520 cfs (15 ems), it was one of the largest in the <br />upper Colorado River basin (Follansbee, 1929). Larger and more <br />significant impacts related to construction of dams continued throughout <br />the 20th century. These projects have had a significant impact on the <br />character of the Colorado River by changing its natural flow regime, <br />turbidity, temperature, and aquatic community (Fradkin, 1981). In <br />conjunction with these impacts, the introduction of nonnative vegetation <br />and fish species has placed additional stress on native fish communities <br />(Carlson and Muth, 1989). Changes in flow regime and negative <br />interactions with nonnative fishes are believed to be the primary causes of <br />the severe decline in their populations (Tyus, 1992). There are currently <br />four federally-listed endangered fishes in the upper Colorado River basin: <br />the Colorado squaw fish (Ptychocheillts Iucius), razorback sucker <br />