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<br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br /> <br />densi ty nursery areas for the species (Biological Sub-Conuni ttee 1984). The <br />reader is reminded that the CPE's in this report are the result of sampling <br />for larvae and YOY throughout the summer, in contrast to the FWS ISMP which <br />assesses CPE for YOY squawfish only from sampling in the fall, September or <br />October. As a result, the CPE's in this report are probably higher than com- <br />parable CPE's from the standardized monitoring program for the respective <br />regions. <br /> <br />ii. Hatching Dates. In order to gain a better understanding of the <br />envi ronmental factors that influence hatching success and survival of YOY <br />Colorado squawfish, hatching dates for 490 YOY and larvae were back-calculated <br />using the length/age regressions developed by Haynes et al. (1985). The <br />relationships are as follows: <br /> <br />REGRESSION A: <br /> <br />For fish with a total length of less than 22 rom <br /> <br />Age = -76.7105 + 17.4949L - 1.055L2 + 0.0221L3 (r2 = 0.99) <br /> <br />REGRESSION B: <br /> <br />For fish with total lengths of 22.0 to 47.0 rom <br /> <br />Age = -26.6421 + 2.7798L (r2 = 0.99) <br /> <br />When the back-calculated dates of hatching are plotted on a frequency histogram <br />(Figure 3), it becomes evident that in 1987, hatching of Colorado squawfish, <br />captured in the Cataract Canyon region, occurred primarily from mid-June <br />through late July. It appears from this analysis that some fish also hatched <br />during May. This histogram represents hatching times very different from those <br />seen in 1986, when hatching peaked much later; in early August. When compared <br />with a similar histogram developed for YOY captured from the same area in 1985 <br />(Valdez 1985), differences in hatching times between the three years become <br />evident. In 1985, hatching began the first of June, peaked during the first <br />week of July, and continued through the first week of August. Clearly, <br />hatching of Colorado squawfish captured in Cataract Canyon occurred at dif- <br />ferent times of the summer for the 3 years examined. Spawning dates can be <br />determined by subtracting 3-5 days (Hamman, 1981) from the hatching dates. <br /> <br />Some of the variation seen in back-calculated hatching dates might be <br />attributed to differential growth rates, slight errors in length measurements <br />afield, or spawning events separated by time. But, the most likely explanation <br />for this variation is that since Cataract Canyon is located below the con- <br />fluence of the Green and Colorado rivers, it receives recently-hatched fish <br />being transported from more than one spawning site upstream on ei ther river. <br />It is also possible that some of these fish originated from spawning sites in <br />or near Cataract Canyon. <br /> <br />In an effort to assess the effect of the two variables, water temperature and <br />river flow, on the hatching times of Colorado squawfish, these were plotted <br />wi th the frequency histograms in Figure 3. Temperature data for 1985 and 1986 <br />are from the bottom readings of the BOR gauge at Clearwater Canyon in Lake <br />Powell (about 8 miles below Imperial Canyon). Temperatures dates in 1987 is <br /> <br />19 <br />