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<br />3.0 Scientific Basis And Underlying Principles <br /> <br />3-15 <br /> <br />April 2004 <br /> <br />The geomorphic and hydrologic characteristics of given floodplains that maximize larval <br />entrainment are not well understood. It is assumed that drifting larvae are randomly mixed in the <br />river water column and that those floodplains that receive the greatest water volume entrain the <br />greatest numbers of larvae. Studies of drifting surrogate species and artificial beads indicate that <br />these assumptions may not be correct. Numbers of drifting larvae of flannelmouth sucker and <br />bluehead sucker in the Upper Colorado River were greater along shorelines than in the central <br />channel (Valdez et al. 1985). Preliminary studies with artificial beads also indicate that particle <br />distribution may not be random (Personal communication, Kevin Christopherson, Utah Division <br />of Wildlife Resources), and that larval entrainment at a given floodplain may be a function of <br />local geomorphic features (e.g., sand bars, position of floodplain in river bend, number and <br />position of levee openings) and river hydraulics (e.g., local currents, diel river surges). Studies <br />may be necessary to better understand drift and entrainment characteristics of larvae in order to <br />better design floodplain sites. <br /> <br />3.7.3 Sufficient Food Production <br /> <br />Most floodplains produce an abundance of food for fish in the first few months of <br />inundation, although the amount of food produced may vary with floodplain site (Crowl et al. <br />2002; Gourley and Crowl 2002). Timing of inundation and chronology of food production is <br />critical to growth and survival of entrained larvae. Production in floodplains occurs as a <br />chronology of communities that begins with inundation of dry floodplains and the appearance of <br />rich detrital loads, diatoms, and algae. This is followed by emergence of various zooplanktors, <br />such as rotifers and copepods, that transition into larger forms including cladocerans and various <br />insect larvae (Mabey and Shiozawa 1993; Modde 1997; Crowl et al. 2002; Gourley and Crowl <br />2002). Rich detritus and invertebrates are important food sources for young fish (Papoulias and <br />Minckley 1990, 1992), and the timing of their appearance with the entrainment of larvae in these <br />floodplains is critical to larval survival (Wydoski and Wick 1998). Larval razorback sucker pass <br />through a "critical period" when nutrition shifts from endogenous (yolk) to exogenous <br />(zooplankton) sources at between 8 and 19 days of age, and they require immediate sources of <br />moderate to high food densities to avoid starvation (Papoulias and Minckley 1990, 1992; see <br />section 3.5). <br /> <br />3.7.4 Suitable Quantity and Quality Of Water <br /> <br />Depression floodplains must have sufficient depth to maintain suitable water quantity and <br />quality for fish to survive during hot summer days and cold winters for at least 1 year. Some <br />depression floodplains may be perched (i.e., elevation higher than the river bed) and maintaining <br />water in these will require excavation to offset evaporative losses, high water temperatures, low <br />oxygen, and complete ice formation in winter. Other depression floodplains may receive surface <br />inflow or seepage that will help to freshen water quality, moderate temperatures, and prevent <br />total freezing. Suitable water quality in these floodplains is critical to insure maximum fish <br />growth and survival. <br />