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<br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br /> <br />automated output variables. Where possible, critical variables were left as user-specified input <br />variables to provide user-discretion and as much flexibility to the model as possible. A <br />description oflife history variables used in the model is provided in Section 3.0, and the model is <br />described in Section 4.0. Sensitivity analyses were performed to ensure that appropriate input <br />variables were used and to provide a better understanding of the inter-relationships of model <br />variables. <br /> <br />2.2 Task Descriptions <br />Objective 1: Estimate egg production and larval emergence of razorback sucker <br />from spawning sites. <br />1bis objective was addressed by specifying the number of adult razorback sucker in the <br />population, the sex ratio to compute number of females, average size of females to compute <br />weight, number of eggs produced, and survival of eggs to hatching and larvae to emergence. The <br />number of adults, sex ratio, and survival of eggs and larvae are all user-specified input variables. <br />The output of this portion of the model is the total number of larvae' emerging from the spawning <br />bed(s) and available for downstream transport by river currents. <br />Objective 2: Estimate downstream drift of larval razorback sucker and <br />entrainment into floodplain habitats. <br />An important aspect of the survival of razorback sucker is the occurrence of floods and <br />availability of floodplain habitat simultaneous to the emergence and drift of larvae (Wick 1997; <br />Modde and Wick 1997; Muth et al. 1998; Wydoski and Wick 1998). Downstream drift rates and <br />entrainment of larvae into floodplain habitats is an important consideration. The proportions of <br />larval razorback sucker that enter floodplain habitats at varying distances downstream from <br />spawning sites is estimated in the model with the aid of the sub-model "Drift" that allows the <br />user to specify the initial number of drifting larvae (output from Objective 1), mainstem survival <br />rate during drift, and percentage entrainment at specified river miles. The last input variable is <br />provided in order for the user to simulate a particular reach of river; this aspect is illustrated in <br />the sample run of the model described in Section 4.0. <br />Drift time and distance for pelagic eggs and larvae have been simulated using artificial <br />eggs in smaller rivers (Dudley and Platania 1999a, 1999b). Similar approaches were used by <br /> <br />4 <br />