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<br />INTRODUCTION <br /> <br />General <br /> <br />Historically. upper Colorado River basin floodpl ains were inundated <br />annually by flows during spring runoff. but today floodplains are not regularly <br />connected to the river because of channelization by levees. dikes. or rip-rap <br />near population centers and in agricultural areas. Introduced salt cedar or <br />tamarisk (Tamarix spp.) has become establ ished along the shorel ine of major <br />rivers resulting in sediment deposition and stabilized banks that further reduce <br />the connectivity of the river with historic flooded bottomlands (Graf 1978). The <br />periodicity of out.of.channel flooding in the upper Colorado River has <br />dramatically decreased following the onset of transmountain water diversions. <br />irrigation diversions, and the construction of mainstem dams (Osmundson and <br />Kaeding 1991). The construction of dikes coupled with the reduction of high <br />spring flows has altered the natural hydrograph and ei ther reduced or e 1 i mi nated <br />regular flooding of bottomlands. It is believed that flooded bottomlands may <br />have served as nursery areas for the razorback suckerl (McAda 1977: Osmundson and <br />Kaedi ng 1991). In addi t ion, Osmundson and Kaedi ng suggested that oxbow 1 akes and <br />flooded pastures in the Grand Valley near Grand Junction. Colorado, were <br />historically the primary spawning habitats of the razorback sucker. They also <br />suggested that low velocity. off-channel habitats were used by adult Colorado <br />squawfish during high streamflows. <br /> <br />Numerous studies (Grawboski and Hiebert 1989; Tyus and Karp 1989; Wydoski <br />and Wick 1994) have suggested the importance of seasonal flooding to river <br />product i vi ty . Other studi es have suggested that flooded bottom 1 ands were <br />important to adult razorback sucker for feeding prior to and after spawning and <br />for the rearing of their young (Tyus and Karp 1989). Adult fish may have used <br />these off. channel habitats for "velocity shelters" to escape the high water <br />velocities from the spring runoff. <br /> <br />The turbid rivers in the upper basin are not very productive for <br />zooplankton that are essential for survival during the early life s~ages of the <br />razorback sucker. When compared to the riverine environment! and river <br />backwaters, inundated bottomlands produce the highest densities of [zooplankton <br />(Grabowski and Hiebert 1989: Mabey and Shiozawa 1993). Although pnedation has <br />been documented to be a limiting factor in survival of larval razo~back sucker <br />in the lower basin (Minckley et al. 1991). starvation may also li~it survival <br />(Marsh and Langhorst 1988; Papoul ias and Minckley 1990). It is hypot~esized that <br />the loss of these productive flooded bottomland habitats appears tojbe limiting <br />the recruitment of the razorback sucker in the upper Colorado River b~sin because <br />of the lack of the right sizes of food organisms at the right time and in <br />sufficient quantity (Wydoski and Wick 1994). <br /> <br />Flooded bottoml and habi tats occur in broad vall eys along l!ow grad; ent <br />stream reaches. Bottomlands are off. or out-of-channel habitats that include <br /> <br />1 Scientific names and two-letter codes of all fishes mentioned in this report are give~ in Appendix A: <br />Table A.I. Only common names for these fishes are used in the text. ' <br />