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<br />I <br />I <br /> <br />Only 43% of 250 fish control projects were considered to be successful in <br />a comprehensive review by Meronek et al. (1996). Total control of <br />nonnative fishes is impossible in the Upper Colorado River Basin because <br />these fishes are well established with self-sustaining populations (Wiley <br />and Wydoski 1993). However, attempts to remove nonnative fishes should <br />help to increase the numbers of razorback suckers in Upper Basin rivers. <br />Minckley and Meffe (1987) believed that even temporary removal or <br />suppression of nonnative predators or competitors may enhance native fish <br />populations. Although biologists are often intimidated by the <br />inefficient and labor-intensive methods of fish control and possible <br />negative public reaction, there is a necessity to reduce (or eradicate if <br />possible) nonnative fish species because of the magnitude of the problem <br />in certain waters (Temple 1990). Ultimately, the endangered fishes must <br />be able to sustain their populations with the established nonnative <br />fishes in the Upper Basin, if recovery is to be realized. Otherwise, <br />periodic or continuous human intervention will be required to control <br />nonnative, warmwater fishes because compensatory growth and survival of <br />nonnative fishes will allow rapid resiliency (Wiley and Wydoski 1993). <br /> <br />I <br />I <br />I <br />I <br />f <br />I <br />l <br />It <br /> <br />t <br /> <br />Mechanical control methods are most practical for management of nonnative <br />fishes in the Upper Colorado River Basin. Chemical control in the main <br />channels or connected habitats would be undesirable based on numerous <br />accidental fish kills that have occurred in running waters. Chemical <br />control is a viable option for control of nonnative fishes in floodplain <br />ponds and is being implemented to reduce chronic escapement from gravel- <br />pit ponds along the upper Colorado and Gunnison rivers (U.S. Fish and <br />wildlife Service 1998). Biological control generally uses predatory fish <br />species that will consume any suitable fish species including endangered <br />fishes. Increasing the numbers of Colorado squawfish in the Upper Basin <br />would provide some biological control of nonnative fishes. The use of <br />other fish species as predators and the use of pathogens (e.g., a virus <br />specific to channel catfish) are not viable options because of high risk. <br />Partial control using mechanical control methods (e.g., removal with <br />various gear, increased water velocity, increased streamflows, etc.) is <br />the only option that reduces risks but this control method often does not <br />generally remove an adequate proportion of the nonnative fish population <br />and compensatory mechanisms of increased growth and fecundity will allow <br />rapid repopulation (i.e., resiliency) by nonnative fishes (Wiley and <br />Wydoski 1993). Osmundson and Kaeding (1991) suggested that streamflow <br />manipulations might be used to manage nonnative fishes while enhancing <br />native fishes. 'However, Valdez (1990) concluded that regulation of <br />streamflows may not be an effective long-term method to reduce nonnative <br />fishes that are adapted to river environments or with a high reproductive <br />potential. Valdez reported that red shiners were reduced in numbers <br />during a year with high streamflows in Cataract Canyon of the Colorado <br />River but exhibited a high resiliency during the following year with a <br />lower streamflow. Partial control of nonnative fishes should be <br />evaluated in river reaches where experimental floodplain enhancement or <br />restoration is completed to determine the responses of native (including <br />endangered) and nonnative fishes. All habitat enhancement or restoration <br />endeavors and nonnative fish control measures should be implemented <br />concurrently by applying adaptive management (Walters 1986; Walters and <br />Hillborn 1978) that allows actions to be taken even when there is a great <br />deal of uncertainty (Ludwig et al. 1993). <br /> <br />t <br /> <br />t <br /> <br />, <br /> <br />~ <br />t <br /> <br />~ <br /> <br />Cover requirements for fish varies by species and life stage that may <br />differ diurnally and seasonally (Gore and Shields 1995). Cover reduction <br />in river ecosystems may reduce fish populations up to 80% (Wesche 1985). <br />Mortality of Age-O razorback suckers may be less in depression ponds with <br />rooted aquatic vegetation that serves as escape cover along the middle <br />Green River (Modde 1997). However, there is no escape cover in gravel- <br />pit ponds without rooted aquatic vegetation along the upper Colorado <br /> <br />35 <br />