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<br />002482 <br /> <br />Narratives For Biological Resources <br />Kanab Ambersnail <br /> <br />Distribution and Abundance <br /> <br />t', <br /> <br />Kanab ambersnail (KAS; Succineidae: Oxvloma havdeni kanabensis Pilsbry 1948), is a landsnail <br />that was proposed for emergency listing (U.S. Fish and Wildlife Service 1991a, 1991b) and <br />officially listed and endangered in 1992 (U.S. Fish and Wildlife Service 1992). Fossil Oxvloma <br />shells have been recovered from sediments in Grand Gulch (lower San Juan River) that date to <br />9,200 years ago (Kerns 1993). Living KAS were fust collected by J.H. Ferriss in 1909 near <br />Kanab, Utah in seep vegetation (Ferriss 1910, Pilsbry and Ferriss 1911, Pilsbry 1948). Extant <br />populations of KAS are presently known to occur at two southwestern springs: one at Three <br />Lakes, near Kanab Utah, and the other at Vaseys Paradise, a spring at Colorado River Mile <br />31.5R, in Grand Canyon, Arizona (Spamer and Bogan 1993a, 1993b). Two popuiations formerly <br />occurred in the Kanab area, but one population was extirpated by desiccation of its habitat. The <br />remaining Utah population at Three Lakes occurs at several, small spring-fed ponds on cattail <br />(Tvpha sp.; Clarke 1991). The Three Lakes site is privately owned and the land owner is <br />commercially developing the property. <br /> <br />KAS wlire fust collected at Vaseys Paradise in 1991 (Blinn et al. 1992, Spamer and Bogan <br />1993), and an interagency team lead by the Bureau of Reclamation examined KAS ecology there <br />from 1995 through 1997 (Kanab Ambersnail Interagency Work Group 1997a). Vaseys Paradise <br />is a popplar water source and attraction site for Colorado River rafters; however, the dense cover <br />of poiso.n ivy (Toxicodendron rydbergii), and the nearly vertical terrain (Stevens et al. I 997b ) <br />limit access. Within Grand Canyon, KAS is apparently restricted to Vaseys Paradise: more than <br />100 other Grand Canyon springs surveyed from 1991 through 1997 failed to detected the <br />presence of KAS (Stevens, pers. comm., Sorensen and Kubly 1997). Rematched historic <br />photographs ofVaseys Paradise (e.g. Turner and Karpiscak 1980:58-59) reveal that vegetative <br />cover has increased greatly at lower stage elevations since completion of Glen Canyon Dam. <br />Flow regulation by the dam has increased primary KAS habitat area at Vaseys Paradise by more <br />than 40%. <br /> <br />.. <br /> <br />~ <br /> <br />Stevens et aI. (1997b) defined primary KAS habitat at Vaseys Paradise as that dominated by <br />crimson monkeyflower (Mimulus cardinalis), non-native watercress (Nasturtium officinale), <br />sedge (Carex aQuatilis) and smartweed (Polvgonum amphibium). Secondary, or marginal, habitat <br />has been defined as patches of other riparian vegetation that are not dominated by these species <br />and are not used extensively by KAS. Land surveys from 1995 through 1997 revealed rapid <br />changes in vegetation cover over the growing season, with 11.2% to 16.1 % of the estimated total <br />primary habitat occurring below the 45,000 cfs stage in 1995, and 7.0-12.0% of the estimated <br />total habitat occurring from 20 April 1996 through 3 October 1997. The total estimated area of <br />primary habitat was 905.7 m2 (0.22 acres), equivalent to the area of secondary habitat, and the <br />total vegetated area was 1811.4 m2 (0.44 acres) in June, 1995. <br /> <br />Life Requisites <br /> <br />KAS occurs on little-disturbed, saturated soil and associated wetland vegetation at Three Lakes, <br />near Kanab, Utah (Stevens, pers. commun.), where cattail and sedges are the predominant <br />